Peer-Reviewed Publications

41.    Clark, C. J., and Mistick, E. A.  Accepted.  Humming hummingbirds, insect flight tones, and a model of animal flight sound. Journal of Experimental Biology. DOI: 10.1242/jeb.214965


 40. Valentina Gómez-Bahamón, Diego T Tuero, María Isabel Castaño, Alex E Jahn, John M. Bates, C. J. Clark. 2020. Sonations in Migratory and Non-migratory Fork-tailed Flycatchers (Tyrannus savana). Integrative Comparative Biology. DOI: 10.1093/icb/icaa115     .pdf


 39.  LePiane, Krista and Clark, C. J. 2020. The dorsal velvet of Barn Owl wing feathers decreases rubbing sounds during flapping flight. Integrative and Comparative Biology. DOI:10.1093/icb/icaa045 .pdf


 38.   Niese, Robert L, Clark, C. J. and Tobalske, Bret W. 2020. Specialized feathers produce sonations during flight in Columbina ground doves. Integrative and Comparative Biology. doi: 10.1093/icb/icaa051 .pdf


 37. Clark, C. J., LePiane, Krista and Liu, Lori. 2020. Evolutionary and ecological correlates of quiet flight in nightbirds, hawks, falcons, and owls. Integrative Comparative Biology. doi.org/10.1093/icb/icaa039   .pdf


 36. Johnson, K. E., Clark, C. J. 2020. Ontogeny of vocal learning in a hummingbird. Animal Behavior. 167, 139-150. DOI: 10.1016/j.anbehav.2020.07.010   .pdf


 35. Clark, C. J., Le Piane, K, and Liu, L. 2020. Evolution and ecology of silent flight in owls and other flying vertebrates. Integrative Organismal Biology. 2: 1-32. DOI: 10.1093/iob/obaa001  .pdf

34. Clark, C. J. and Rankin, D. 2020. Subtle, pervasive genetic correlation between the sexes in the evolution of dimorphic hummingbird tail sexual ornaments. Evolution. https://doi.org/10.1111/evo.13881         .pdf

33. Myers, B. M. Rankin, D., Burns, K., and Clark, C. J.  2019. Behavioral and morphological evidence of an Allen’s × Rufous Hummingbird (Selasphorus sasin × S. rufus)             hybrid zone in southern Oregon and northern California. The Auk: Ornithological Advances, pp 1-24  DOI: 10.1093/auk/ukz049 pdf

  1. Clark, C. J., Rankin, D, and Johnson, K. 2018 Female song in Costa's Hummingbird (Calypte costae). Wilson J. of Ornith, 130:987-992 .pdf
  2. Clark, C. J. 2018. Signal or cue? Locomotion-induced sounds and the evolution of communication. Anim. Behav. 143:83-91 . pdf
  3. Clark, C. J., and Mistick, E. A. 2018. Kinematic control of Allen's Hummingbird wing trill over a range of flight speeds. J. Exp. Biol., 221:. DOI: 10.1242/jeb.173625 pdf
  4. Clark, C. J., and Mistick, E. A. 2018. Strategic acoustic control of a hummingbird courtship dive. Current Biology. 28: 1-8. 10.1016/j.cub.2018.03.021 pdf
  5. Rankin, D, Clark, C. J. and Rankin, E. 2018. Hummingbirds use taste and touch to discriminate against nectar resources that contain Argentine ants. Behav. Ecol. Sociobiol. 72:44. DOI: 10.1007/s00265-018-2456-z pdf
  6. Clark, C. J., McGuire, J A. Bonaccorso, E, Berv, J. S. Prum, R. O. 2018. Complex coevolution of wing, tail, and vocal sounds of courting male bee hummingbirds. Evolution, 72: 630-646. pdf
  7. Clark, C. J., Mountcastle, A. M., Mistick, E. A., Elias, D. O. 2017. Resonance frequencies of honeybee (Apis mellifera) wings. J. Exp. Biol., 220: 2697-2700. DOI: 10.1242/jeb.154609. pdf
  8. Clark, C. J. 2017. Bird records show substantial growth of the Allen's Hummingbird (Selasphorus sasin sedentarius) population in urban Southern California. The Condor: Ornithological Applications. 119: 122-130. doi: 10.1650/CONDOR-16-153.1 pdf
  9. Clark, C. J., Kirschel, A. N. G., Hadjioannou, L., and Prum, R. O. 2016. Smithornis broadbills produce loud wing song by aeroelastic flutter of inner primary wing feathers. J. Exp. Biol., 219: 1069-1075.  pdf
  10. Clark, C. J. and Prum, R. O. 2015. Aeroelastic flutter of feathers, flight, and the evolution of nonvocal communication in birds. J. Exp. Biol. 218: 3520-3527. doi:10.1242/jeb.126458  pdf   .... Table S1 with post-publication updates
  11. Feo, T. J., Musser, J., Berv, J. and Clark, C. J. 2015. Divergence in morphology, calls, song, mechanical sounds, and genetics supports species status for the Inaguan hummingbird (Trochilidae: Calliphlox "evelynae" lyrura). Auk, 132: 248-264. 10.1642/AUK-14-108.1 pdf
  12. Clark, C. J. 2014. Harmonic hopping, and both punctuated and gradual evolution of acoustic characters in Selasphorus hummingbird tail feathers. PLOS ONE, 9: 10.1371/journal.pone.0093829
  13. Clark, C. J., Elias, D. O., Girard, M. B. and Prum, R. O. 2013. Structural resonance and mode of flutter of hummingbird tail feathers. J. Exp Biol., 216: 3404-3413. pdf
  14. Clark, C. J., Elias, D. O., and Prum, R. O. 2013. Hummingbird feather sounds are produced by aeroelastic flutter, not vortex-induced vibration. J. Exp Biol., 216: 3395-3403. pdf
  15. Clark, C. J., Feo, T. J. and van Dongen, W. 2013. Sounds and courtship displays of the Peruvian Sheartail, Chilean Woodstar, Oasis Hummingbird, and a hybrid male Peruvian Sheartail × Chilean Woodstar. Condor, 115: 560-577. pdf
  16. Clark, C. J. 2012. The role of power versus energy in courtship: what is the “energetic cost” of a courtship display? Anim. Behav., 84: 269-277. pdf
  17. Clark, C. J., Feo, T. J., and Bryan, K. B. 2012. Courtship displays and sonations of a male hybrid Broad-tailed × Black-chinned Hummingbird (Selasphorus platycercus × Archilochus alexandri). Condor, 114: 329-340. pdf
  18. Clark, C. J., Elias, D., and Prum, R. O. 2011. Aeroelastic flutter produces hummingbird feather songs. Science, 333: 1430-1433. pdf
  19. Clark, C. J. 2011. Wing, tail, and vocal contributions to the complex signals of a courting Calliope Hummingbird. Curr. Zool, 57: 187-196. pdf
  20. Clark, C. J., Feo, T. J. and Escalante, I. 2011. Courtship displays and natural history of the Scintillant (Selasphorus scintilla) and Volcano (S. flammula) hummingbirds. Wilson J. Ornith., 123: 218-228. pdf
  21. Feo, T. J. and Clark, C. J. 2010. The displays and mechanical sounds of the Black-chinned Hummingbird (Trochilidae: Archilochus alexandri). Auk, 127: 787-796. pdf
  22. Clark, C. J. 2010. Effects of tail length on an escape maneuver of the Red-billed Streamertail. J. Ornith., 152: 397-408. DOI: 10.1007/s10336-010-0550-8. pdf
  23. Clark, C. J. and Dudley, R. 2010. Hovering and forward flight energetics in Anna’s and Allen’s Hummingbirds. Physiol. Biochem. Zool. 83: 654-662. DOI: 10.1086/653477 pdf
  24. Brennan, P. L. R., Clark, C. J. and Prum, R. O. 2010. Explosive eversion and functional morphology of the waterfowl penis supports sexual conflict in genitalia. Proc. R. Soc. Lond. B, 277: 1309-1314. DOI: 10.1098/rspb.2009.2139. pdf
  25. Clark, C. J. 2010. The evolution of tail shape in hummingbirds. Auk, 127: 44-56. pdf
  26. Clark, C. J. and Feo, T. J.* 2010. Why do Calypte hummingbirds “sing” with both their tail and their syrinx? An apparent example of sexual sensory bias. Am. Nat., 175: 27-37. pdf
  27. Clark, C. J. 2009. Courtship dives of Anna’s Hummingbird offer insights into flight performance limits. Proc. R. Soc. Lond. B. 276: 3047-3052. pdf
  28. Clark, C. J. and Dudley, R. 2009. Flight costs of long, sexually selected tails in hummingbirds. Proc. R. Soc. Lond. B. 276: 2109-2115. pdf
  29. Clark, C. J. 2008. Fluttering wing feathers produce the flight sounds of male streamertail hummingbirds. Biology Letters, 4: 341-344. pdf
  30. Clark, C. J. and Feo, T. J.* 2008. The Anna’s Hummingbird chirps with its tail: a new mechanism of sonation in birds. Proc. R. Soc. Lond. B. 275: 955-962. pdf
  31. Tobalske, B. W., Warrick, D. R., Clark, C. J., Powers, D. R., Hedrick, T. L., Hyder, G. & Biewener, A. A. 2007. Three-dimensional kinematics of hummingbird flight. J. Exp. Biol., 210: 2368-2382. pdf
  32. Clark, C. J. 2006. Observations on the Vervain Hummingbird’s (Mellisuga minima) display dive and territorial behavior. Ornitologia Neotropical 17: 403-408. pdf

Other Publications

    Clark, C. J., and Jaworski, J. 2020. Introduction to the symposium: Bio-inspiration of quiet flight of owls and other flying animals: recent advances and unanswered questions. Integrative Comparative Biology.   DOI: 10.1093/icb/icaa128    .pdf

  • Clark, C. J. 2016. Locomotion-Induced Sounds and Sonations: Mechanisms, Communication Function, and Relationship with Behavior. In Vertebrate Sound Production and Acoustic Communication (Suthers, R., and Fitch, T., eds.). Springer Handbook of Auditory Research, 53: 83-117.  pdf
  • Clark, C. J. 2013. Peruvian Sheartail (Thaumastura cora Lesson and Garnot). Neotropical Birds Online (T. S. Schulenberg, Ed.). Cornell Lab of Ornithology, Ithaca, NY. Read it here
  • Clark, C. J. and Mitchell, D. 2013. Allen’s Hummingbird (Selasphorus sasin). The Birds of North America # 501 (Alan Poole, Ed.). Cornell Lab of Ornithology, Ithaca, NY. Read it here
  • Clark, C. J., and Russell, S. M. 2012. Anna’s Hummingbird (Calypte anna). The Birds of North America # 226 (Alan Poole, Ed.). Cornell Lab of Ornithology, Ithaca, NY. Read it here
  • Clark, C. J. 2011. Scintillant Hummingbird (Selasphorus scintilla Gould), Neotropical Birds Online (T. S. Schulenberg, Ed.). Ithaca: Cornell Lab of Ornithology, Ithaca, NY. Read it here
  • Clark, C. J. 2009. Vervain Hummingbird (Mellisuga minima Linnaeus), Neotropical Birds Online (T. S. Schulenberg, Ed.). Cornell Lab of Ornithology, Ithaca, NY. Read it here
  • Altshuler, D. L., and Clark, C. J. 2003. Darwin’s Hummingbirds [Perspective]. Science, 300: 588-589. pdf